Gained his Ph.D. in 1984 from the University of Cambridge, working with Sir Martin Evans on the early characterization of embryonic stem cells, later joining Rudolf Jaenisch’s group developing animal models of connective tissue disorders. He has worked with Angelika Schnieke since 1984. From 1992–2003 they worked with the biotechnology company PPL Therapeutics in Edinburgh, Scotland, where Angelika Schnieke gained her Ph.D. from the University of Edinburgh for the development of somatic cell nuclear transfer as a means of generating transgenic sheep. Their current research focuses on animal stem cells and the genetic engineering of livestock for bio-medical applications.
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